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<br /> 3 June 2010 | www.nature.com/nature | $10 THE INTERNATIONAL WEEKLY JOURNAL OF SCIENCE<br /> SPREADING<br /> THE RISK<br /> Population diversity boosts<br /> fishery resilience<br /> UNDER THE VOLCANO Naturejobs<br /> Polar research<br /> An eruption of science<br /> COSMIC EVOLUTION<br /> Beyond the standard model<br /> MATHEMATICAL MENTORS<br /> Protégé performance across<br /> the centuries<br /> $10.00US $12.99CAN<br /> 22<br /> 0 71486 03070 6<br /> Vol 465 | 3 June 2010 | doi:10.1038/nature09060<br /> LETTERS<br /> Population diversity and the portfolio effect in an<br /> exploited species<br /> Daniel E. Schindler1, Ray Hilborn1, Brandon Chasco1, Christopher P. Boatright1, Thomas P. Quinn1, Lauren A. Rogers1<br /> & Michael S. Webster2<br /> One of the most pervasive themes in ecology is that biological diver-<br /> s<a title="68228-elite-nature-june2010.pdf page 1" href="http://viewer.zmags.com/publication/5faee868?page=1"> 3 June 2010 | www.nature.com/nature | $10 </a> <a title="68228-elite-nature-june2010.pdf page 2" href="http://viewer.zmags.com/publication/5faee868?page=2"> Vol 465 | 3 June 2010 | doi:10.1038/nature09060 </a> <a title="68228-elite-nature-june2010.pdf page 3" href="http://viewer.zmags.com/publication/5faee868?page=3"> LETTERS </a> <a title="68228-elite-nature-june2010.pdf page 4" href="http://viewer.zmags.com/publication/5faee868?page=4"> NATURE | Vol 465 | 3 June 2010 </a> <a title="68228-elite-nature-june2010.pdf page 5" href="http://viewer.zmags.com/publication/5faee868?page=5"> LETTERS </a>